With no explanation, label each line with "O", "other-biologything", "location-GPE", "organization-education", "other-educationaldegree", "art-broadcastprogram", "art-film", "art-music", "art-other" or "art-painting" preceded by ":".
A
graduate
of
the
University
of
Natal
(
South
Africa
)
,
with
a
PhD
from
Göteborg
University
(
Sweden
)
,
she
is
best
known
for
her
research
on
the
evolution
of
sexual
signals
in
the
Red-collared
widowbird
and
more
recently
research
on
maternal
effects
and
the
evolution
of
alternative
reproductive
strategies
in
the
Gouldian
finch
.
A:O
graduate:O
of:O
the:O
University:organization-education
of:organization-education
Natal:organization-education
(:O
South:O
Africa:location-GPE
):O
,:O
with:O
a:O
PhD:other-educationaldegree
from:O
Göteborg:organization-education
University:organization-education
(:O
Sweden:location-GPE
):O
,:O
she:O
is:O
best:O
known:O
for:O
her:O
research:O
on:O
the:O
evolution:O
of:O
sexual:O
signals:O
in:O
the:O
Red-collared:other-biologything
widowbird:other-biologything
and:O
more:O
recently:O
research:O
on:O
maternal:O
effects:O
and:O
the:O
evolution:O
of:O
alternative:O
reproductive:O
strategies:O
in:O
the:O
Gouldian:other-biologything
finch:other-biologything
.:O